Author contributions
Y.H. initiated the research idea. K.J.W. and Y.H. discussed with T.R.K. and codesigned the study. C.S.A. and F.V. collected and processed sediment samples. K.J.W., M.M., and N.R. analyzed the DNA
and the phylogenetic data. S.L. and K.J.W. conducted the alkenone
analyses. K.J.W. and Y.H. cowrote the manuscript with contributions from
coauthors.