Acknowledgments
The study was carried out as part of the National Park Service’s Critical Connections Program. All capture, tagging, and sampling work were conducted under permits’ NPS IACUC AKR McIntyre Birds 2015.A3’ and ’AK DENA WRST McIntyre Birds 2018.A3’ and USGS Banding permit #24141. We thank the National Park Service, Alaska Geographic, Denali Education Center, Murie Science and Learning Center, Camp Denali, and Biodiversity Research Institute for funding and/or in-kind support for the Critical Connections Program. We thank everyone who participated in the study, particularly Tucker Grigsby, Carl Malatin, and Jason Reppert, for outstanding field assistance and logistic supply. We also thank Laura Phillips and Emily Williams, who helped with many aspects of the field work and reporting and led the field efforts in 2018.