4 Key Laboratory for Conserving Wildlife with Small Populations in Yunnan, Faculty of Biodiversity Conservation, Southwest Forestry University, Kunming, China.
*Xing-Jin Che and Yuan-Wei Zhang contributed equally to the work.
**Correspondence: Xiao-Ai Wang, State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, The Innovative Academy of Seed Design, Chinese Academy of Sciences, No.17 Longxin Road, Kunming 650201, Yunnan, China. E-mail: wangxiaoai@mail.kiz.ac.cn