1. Introduction
1.1 Importance of animal-mediated seed dispersal
Seed dispersal is one of the most important ecosystem services globally (Aslan et al, 2013). Seed dispersal shapes plant populations by ensuring their regeneration through gene flow and increasing the spacing of individuals, which reduces the likelihood of density-dependent mortality near parental plants (Jansen et al, 2008; Jordano, et al, 2011). Humans indirectly benefit from this global service through the seed dispersal of valuable timber species, and edible and medicinal plants (Wenny et al, 2016), yet seed dispersal is one of the most threatened processes in plant regeneration, worldwide (Neuschulz et al, 2016). Habitat loss and fragmentation are the main threats to seed dispersal as they restrict the movement and natural behaviour of local seed dispersers (Mahoney et al, 2018; Browne and Karubian, 2018). Accurate measurement of seed dispersal distances is therefore essential to fully understand the effect of habitat loss on critical ecosystem services.