References
1. Hadley N. 1994 Water Relations of Terrestrial Arthropods . New
York: Academic Press.
2. Benoit JB, Grosso N, Yoder J, Denlinger DL. 2007 Resistance to
dehydration between bouts of blood feeding in the bed bug, Cimex
lectularius , is enhanced by water conservation, aggregation, and
quiescence. Am J Trop Med Hyg 76 , 987–993.
3. Klok CJ, Chown SL. 1999 Assessing the benefits of aggregation:
thermal biology and water relations of anomalous Emperor Moth
caterpillars. Funct. Ecol. 13 , 417–427.
4. Yoder JA, Grojean NC. 1997 Group influence on water conservation in
the giant Madagascar hissing-cockroach, Gromphadorhina portentosa(Dictyoptera: Blaberidae). Physiol. Entomol. 22 , 79–82.
(doi:10.1111/j.1365-3032.1997.tb01143.x)
5. Johnson R. 2021 Desiccation limits recruitment in the pleometrotic
desert seed-harvester ant Veromessor pergandei . Ecol.
Evol. 11 , 294–308. (doi:10.1002/ece3.7039)
6. Ostwald M, Haney B, Fewell J. 2022 Ecological Drivers of Non-Kin
Cooperation in the Hymenoptera. Front. Ecol. Evol. 10 ,
768392. (doi:doi: 10.3389/fevo.2022.768392)
7. Jetz W, Rubenstein DR. 2011 Environmental Uncertainty and the Global
Biogeography of Cooperative Breeding in Birds. Curr. Biol.21 , 72–78. (doi:10.1016/j.cub.2010.11.075)
8. La Richelière F et al. 2022 Warm and arid regions of the world
are hotspots of superorganism complexity. Proc. R. Soc. B Biol.
Sci. 289 . (doi:10.1098/rspb.2021.1899)
9. Lukas D, Clutton-Brock T. 2017 Climate and the distribution of
cooperative breeding in mammals. R Soc Open Sci 4 ,
160897.
10. Purcell J. 2011 Geographic patterns in the distribution of social
systems in terrestrial arthropods. Biol. Rev. 86 ,
475–491. (doi:10.1111/j.1469-185X.2010.00156.x)
11. Purcell J, Avilés L. 2008 Gradients of precipitation and ant
abundance may contribute to the altitudinal range limit of subsocial
spiders: insights from a transplant experiment. Proc R Soc B275 , 2617–2625. (doi:10.1098/rspb.2008.0582)
12. Bollazzi M, Roces F. 2010 Leaf-cutting ant workers (Acromyrmex
heyeri) trade off nest thermoregulation for humidity control. J.
Ethol. 28 , 399–403. (doi:10.1007/s10164-010-0207-3)
13. Lindauer M. 1954 Temperaturregulierung und Wasserhaushalt im
Bienenstaat. Z Vergl Physiol 36 , 391–432.
14. Nicolson SW. 2009 Water homeostasis in bees, with the emphasis on
sociality. J. Exp. Biol. 212 , 429–434.
(doi:10.1242/jeb.022343)
15. Ostwald M, Smith ML, Seeley TD. 2016 The behavioral regulation of
thirst, water collection and water storage in honey bee colonies.J. Exp. Biol. 219 , 2156–2165. (doi:10.1242/jeb.139824)
16. Bohart G, Youssef N. 1976 The Biology and Behavior of Evylaeus
galpinsiae . Wasmann J. Biol. 34 , 185–234.
17. Danforth B. 1991 Female Foraging and Intranest Behavior of a
Communal Bee, Perdita portalis (Hymenoptera: Andrenidae).Ann. Entomol. Soc. Am. 84 , 537–548.
18. McCorquodale DB. 1989 Soil softness, nest initiation and nest
sharing in the wasp, Cerceris antipodes (Hymenoptera: Sphecidae).Ecol. Entomol. 14 , 191–196.
19. Ostwald M, Fox TP, Hillery WS, Shaffer Z, Harrison JF, Fewell JH.
2022 Group-living carpenter bees conserve heat and body mass better than
solitary individuals in winter. Anim. Behav. 189 ,
59–67. (doi:10.1016/j.anbehav.2022.04.012)
20. Pfennig W. 1995 Absence of joint nesting advantage in desert seed
harvester ants: evidence from a field experiment. Anim Behav49 , 567–575.
21. Berg A, Sheffield J. 2018 Climate Change and Drought: the Soil
Moisture Perspective. Curr. Clim. Change Rep. 4 ,
180–191. (doi:10.1007/s40641-018-0095-0)
22. Blumstein DT, Hayes LD, Pinter-Wollman N. 2022 Social consequences
of rapid environmental change. Trends Ecol. Evol. ,
S0169534722002853. (doi:10.1016/j.tree.2022.11.005)
23. Ostwald MM, da Silva CRB, Seltmann K. 2023 How does climate change
impact social bees and bee sociality? Authorea Preprints(doi:10.22541/au.169175894.41597921/v1)
24. Heinze J, Kellner K, Seal J. 2017 Sociality in Ants. InComparative Social Evolution (eds DR Rubenstein, P Abbot), pp.
21–49. Cambridge: Cambridge University Press.
25. Michener C. 2007 The Bees of the World . Baltimore, MD: Johns
Hopkins University Press.
26. Ross K, Matthews R, editors. 1991 The Social Biology of
Wasps . Ithaca, NY: Comstock Publishing Associates.
27. Clutton-Brock T. 2009 Cooperation between non-kin in animal
societies. Nature 461 , 51–57. (doi:10.1038/nature08366)
28. Hurd P, Linsley E. 1959 Observations on the nest-site behavior ofMelissodes composita Tucker and its Parasites, with notes on the
communal use of nest entrances. Entomol. News LXX ,
141–146.
29. R Core Team. 2023 R: A Language and Environment for Statistical
Computing. R Found. Stat. Comput.
30. Fox J, Weisberg S. 2019 An {R} Companion to Applied
Regression, Third Edition . Thousand Oaks, CA: Sage.
31. Boratynski JS, Willis CKR, Jefimow M, Wojciechowski S. 2015
Comparative Biochemistry and Physiology, Part A Huddling reduces
evaporative water loss in torpid Natterer’s bats, Myotis nattereri.Comp. Biochem. Physiol. Part A 179 , 125–132.
(doi:10.1016/j.cbpa.2014.09.035)
32. Gilbert C, Robertson G, Le Maho Y, Naito Y, Ancel A. 2006 Huddling
behavior in emperor penguins: Dynamics of huddling. Physiol.
Behav. 88 , 479–488. (doi:10.1016/j.physbeh.2006.04.024)
33. Jones JC, Oldroyd BP. 2006 Nest Thermoregulation in Social Insects.
In Advances in Insect Physiology , pp. 153–191. Elsevier.
(doi:10.1016/S0065-2806(06)33003-2)
34. Krause J, Ruxton GD. 2002 Living in Groups . Oxford: Oxford
University Press.
35. Yahav S, Buffenstein R. 1991 Huddling Behavior Facilitates
Homeothermy in the Naked Mole Rat Heterocephalus glaber. Physiol.
Zool. 64 , 871–884.
36. Bong L-J, Neoh K-B, Yoshimura T. 2018 Comparison of Water Relation
in Two Powderpost Beetles Relative to Body Size and Ontogenetic and
Behavioral Traits. Environ. Entomol. 47 , 990–996.
(doi:10.1093/ee/nvy062)
37. Rasa OAE. 1997 Aggregation in a Desert Tenebrionid Beetle: A
Cost/Benefit Analysis. Ethology 103 , 466–487.
(doi:10.1111/j.1439-0310.1997.tb00161.x)
38. Yoder JA, Denlinger DL, Wolda H. 1992 Aggregation promotes water
conservation during diapause in the tropical fungus beetle, Stenotarsus
rotundus. Entomol. Exp. Appl. 63 , 203–205.
39. Broly P, Devigne L, Deneubourg J-L, Devigne C. 2014 Effects of group
size on aggregation against desiccation in woodlice (Isopoda:
Oniscidea). Physiol. Entomol. 39 , 165–171.
(doi:10.1111/phen.12060)
40. Derhé M, Moss A, Edwards D, Carmenta R, Hassall M. 2010 Predicting
the effect of climate change on aggregation behaviour in four species of
terrestrial isopods. Behaviour 147 , 151–164.
(doi:10.1163/000579509X12512861455834)
41. Willmer PG. 1980 The effects of a fluctuating environment on the
water relations of larval Lepidoptera. Ecol. Entomol. 5 ,
271–292. (doi:10.1111/j.1365-2311.1980.tb01150.x)
42. Southwick EE, Roubik DW, Williams JM. 1990 Comparative energy
balance in groups of africanized and european honey bees: Ecological
implications. Comp. Biochem. Physiol. A Physiol. 97 ,
1–7. (doi:10.1016/0300-9629(90)90713-3)
43. Addo-bediako A, Chown SL, Gaston KJ. 2001 Revisiting water loss in
insects: a large scale view. J. Insect Physiol. 47 ,
1377–1388.
44. Antoine CM, Forrest JRK. 2021 Nesting habitat of ground-nesting
bees: a review. Ecol. Entomol. 46 , 143–159.
(doi:10.1111/een.12986)
45. Cane JH. 1991 Soils of Ground-Nesting Bees (Hymenoptera: Apoidea):
Texture, Moisture, Cell Depth and Climate. J. Kans. Entomol. Soc.64 , 406–413.
46. Lighton JRB, Bartholomew GA. 1988 Standard energy metabolism of a
desert harvester ant, Pogonomyrmex rugosus : Effects of
temperature, body mass, group size, and humidity. Proc. Natl.
Acad. Sci. 85 , 4765–4769. (doi:10.1073/pnas.85.13.4765)
47. McConnell-Garner J, Kukuk PF. 1997 Behavioral Interactions of Two
Solitary, Halictine Bees with Comparisons among Solitary, Communal and
Eusocial Species. Ethology 103 , 19–32.
(doi:10.1111/j.1439-0310.1997.tb00003.x)
48. Packer L. 2006 Use of artificial arenas to predict the social
organisation of halictine bees: Data for fourteen species from Chile.53 , 307–315. (doi:10.1007/s00040-006-0873-x)