Authors’ contribution
Hailei Zheng, Wenjun Hu and Guoxin Shen supervised the overall study, designed the experiment and revised the manuscript. Wenjun Hu and Jiyun Liu analyzed all data and wrote the manuscript. Wenjun Hu and Tingwu Liu performed the proteomic experiments. Lin Chen, Hongling Lu and Chenkai Jiang conducted the physiological and growth indexes determination. Chunquan Zhu and Qian Wu performed the relative gene expression analysis. Tingwu Liu and Chenkai Jiang carried out plant material cultivation and treatment, and experimental material collection.