LT, LA, SLA, LDMC, LC, LN, LP, LN:LP and LC:LN represent leaf thickness,
leaf area, specific leaf area, leaf dry matter content, leaf carbon
content, leaf nitrogen content, leaf phosphorus content, leaf
nitrogen-phosphorus ratio, and leaf carbon-nitrogen ratio, respectively.
*** p < 0.001; ** p < 0.01; * p< 0.05