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Acknowledgements: The authors would like to thank Isabel
Ferreira for her help during fieldwork. Work supported by National Funds
through FCT-Fundação para a Ciência e a Tecnologia in the scope of the
project UIDB/50027/2020. This work was also partially funded by the
European Regional Development Fund (ERDF) through the COMPETE program
and by National Funds through FCT - Foundation for Science and
Technology (project PTDC/BIA-MIC/27995/2017 POCI01-0145- FEDER-027995);
RX was supported by FCT under the Programa Operacional Potencial Humano
– Quadro de Referência Estratégico Nacional funds from the European
Social Fund and Portuguese Ministério da Educação e Ciência
(2020.00854.CEECIND/CP1601/CT0001), AP was supported by FCT
(SFRH/BD/144928/2019).
Data Accessibility: Data will be deposited in NCBI’s Short Read
Archive (SRA) upon acceptance.
Competing Interests: The authors have no competing interests.
Author Contributions: All authors contributed to the writing of
the manuscript; DV collected samples, performed laboratory work and
analyzed the data; DJH conceived the work, collected samples and
contributed with funding; ID-M collected samples; AP performed
laboratory work; RX conceived the work, supervised data analysis and
contributed with funding.
Ethics approval: Experimental protocols and research were
approved by the Portuguese Institute for Conservation of Nature and
Forests (ICNF) (License 703/2021/CAPT).