Author contributions

D.P., P.M., M.S., H.Z. and A.W. designed the study. M.S., H.Z., D.P. and P.M. collected the eDNA samples in the field, with the participation of M.L., C.N., L.P., B.C.S., E.S. and E.T. T.D., J.P. and A.V. conducted the laboratory and bioinformatics analyses. V.B. analysed the data related to the conventional sampling survey. D.P. assisted with data analysis, prepared the figures, and wrote most of the manuscript with significant contributions from the other authors.