ACKNOWLEDEMENTS
We thank the many people who assisted us in field sampling and data collection. We are grateful to the Meigu Dafengding Nature Reserve and Heizhugou Nature Reserve for their permission to do the research. This work received funding from three organizations: a) The investigation on vertbrate in the Meigu Dafengding Nature Reserves no. 16H0859, b) The investigation on vertbrate in the Heizhugou Nature Reserve no. 17H0757 and c) the National Natural Science Foundation of China no 31970495.