Author contributions

M.N., M.G. and B.D. conceived and planned the study. A.I.G., M.G. and M.N. performed the analyses. M.G., B.D., G.W., S.E., A.V. and B.S. contributed to sample preparation and pooled sequence analysis. A.I.G., M.G., B.D. and M.N. contributed to the interpretation of the results. A.I.G. and M.N. took the lead in writing the manuscript. All authors provided critical feedback and helped shape the research, analysis and manuscript.