ACKNOWLEGEMENTS
This study was supported by the National Natural Science Foundation of China (no. 31901109; no. 31901220), GDAS’ Project of Science and Technology Development (2019GDASYL-0105044; 2021GDASYL-20210103056), GDAS Special Project of Science and Technology Development (2018GDASCX-0107). We gratitude to the Forestry Department of Tibet Autonomous Region and Cuona Forestry Bureau for the permits to conduct the researches. We thank Wen Sun for helping us collect data from the field, and we also thank Jianchao Liang for his advice on collecting environmental data.