Conflict of interest 
The authors declare that there is no conflict of interest.
Author contributions : A.P. collected samples, designed and performed the experiments, collected and analyzed the data, and wrote the manuscript; G.W. designed and performed the experiments, C. L. performed the experiments, P.S., N.F., B.G. and F.D provided conceptual advice and edited the manuscript; C.B. designed the study, collected samples, performed the experiments, analyzed the data, and wrote the manuscript.