Acknowledgement
The authors would like to thank Xiao Yang and Juan Wei for their excellent technical assistance. This work was supported by the National Key Research and Development Project (2018YFC0840400), the Gansu major science and technology project (20ZD7NA006), the Collaborative Innovation Project of Chinese Academy of Agricultural Sciences (CAAS-ZDRW202006), the Guangdong Innovative and Entrepreneurial Research Team Program (No. 2014ZT05S123) and the Shenzhen Science and Technology Innovation Commission (JSGG20180508152235868).