REFERENCES
Araújo, M.S., Bolnick, D.I. & Layman, C.A. (2011). The ecological
causes of individual specialisation. Ecol. Lett. , 14, 948–958.
Bastille-Rousseau, G., Rayl, N.D., Ellington, E.H., Schaefer, J.A.,
Peers, M.J.L., Mumma, M.A., et al. (2016). Temporal variation in
habitat use, co-occurrence, and risk among generalist predators and a
shared prey. Can. J. Zool. , 94, 191–198.
Bastille-Rousseau, G., Schaefer, J.A., Mahoney, S.P. & Murray, D.L.
(2013). Population decline in semi-migratory caribou (Rangifer
tarandus ): intrinsic or extrinsic drivers? Can. J. Zool. , 91,
820–828.
Bell, A.M., Hankison, S.J. & Laskowski, K.L. (2009). The repeatability
of behaviour: a meta-analysis. Anim. Behav. , 77, 771–783.
Bjørneraas, K., Van Moorter, B., Rolandsen, C.M. & Herfindal, I.
(2010). Screening global positioning system location data for errors
using animal movement characteristics. J. Wildl. Manage. , 74,
1361–1366.
Bolnick, D.I., Svanbäck, R., Fordyce, J.A., Yang, L.H., Davis, J.M.,
Hulsey, C.D., et al. (2003). The ecology of individuals:
Incidence and implications of individual specialization. Am.
Nat. , 161, 1–28.
Bonar, M., Ellington, E.H., Lewis, K.P. & Vander Wal, E. (2018).
Implementing a novel movement-based approach to inferring parturition
and neonate calf survival. PLoS One , 13, e0192204.
Bonar, M., Lewis, K., Webber, Q.M.R., Dobbin, M., Laforge, M.P. &
Vander Wal, E. (2020). Geometry of the ideal free distribution:
individual behavioural variation and annual reproductive success in
aggregations of a social ungulate. Ecol. Lett. , 23, 1360–1369.
Bonenfant, C., Gaillard, J.M., Coulson, T., Festa-Bianchet, M., Loison,
A., Garel, M., et al. (2009). Empirical evidence of
density-dependence in populations of large herbivores. Adv. Ecol.
Res. , 41, 313–357.
Bradbury, J.W., Vehrencamp, S.L. & Clifton, K.E. (2015). The ideal free
antelope: foraging dispersions. Behav. Ecol. , 26, 1303–1313.
Brashares, J.S., Werner, J.R. & Sinclair, A.R.E. (2010). Social
“meltdown” in the demise of an island endemic: Allee effects and the
Vancouver Island marmot. J. Anim. Ecol. , 79, 965–973.
Cairns, S.J. & Schwager, S.J. (1987). A comparison of association
indices. Anim. Behav. , 35, 1454–1469.
Calenge, C. (2006). The package “adehabitat” for the R software: A
tool for the analysis of space and habitat use by animals. Ecol.
Modell. , 197, 516–519.
Carlson, B.S., Rotics, S., Nathan, R., Wikelski, M. & Getz, W. (2021).
Individual environmental niches in mobile organisms. Nat.
Commun. , 12, 4572.
Charnov, E.L. (1976). Optimal foraging, the marginal value theorem.Theor. Popul. Biol. , 9, 129–136.
DeMars, C.A., Auger-Méthé, M., Schlägel, U.E. & Boutin, S. (2013).
Inferring parturition and neonate survival from movement patterns of
female ungulates: A case study using woodland caribou. Ecol.
Evol. , 3, 4149–4160.
Dingemanse, N.J. & Dochtermann, N.A. (2013). Quantifying individual
variation in behaviour: mixed-effect modelling approaches. J.
Anim. Ecol. , 82, 39–54.
Dingemanse, N.J., Kazem, A.J.N., Réale, D. & Wright, J. (2010).
Behavioural reaction norms: animal personality meets individual
plasticity. Trends Ecol. Evol. , 25, 81–89.
Dochtermann, N.A., Schwab, T. & Sih, A. (2015). The contribution of
additive genetic variation to personality variation: heritability of
personality. Proc. R. Soc. B , 282, 20142201.
Fortin, D., Morris, D.W. & McLouglin, P.D. (2008). Habitat selection
and the evolution of specialists in heterogeneous environments.Isr. J. Ecol. Evol. , 54, 295–309.
Fowler, C.W. (1981). Density dependence as related to life history
strategy. Ecology , 62, 602–610.
Franks, D.W., Weiss, M.N., Silk, M.J., Perryman, R.J.Y. & Croft, D.P.
(2021). Calculating effect sizes in animal social network analysis.Methods Ecol. Evol. , 12, 33–41.
Fretwell, S.D. & Lucas, H.L.J. (1969). On territorial behaviour and
other factors influencing habitat distribution in birds. Acta
Biotheor. , 19, 16–36.
Fussmann, G.F., Loreau, M. & Abrams, P.A. (2007). Eco-evolutionary
dynamics of communities and ecosystems. Funct. Ecol. , 21,
465–477.
Gaillard, J.-M., Festa-bianchet, M., Delorme, D. & Jorgenson, J.
(2000). Body mass and individual fitness in female ungulates: bigger is
not always better. Proc. R. Soc. B , 471–477.
Hadfield, J.D. (2010). MCMC methods for multi-respoinse generalized
linear mixed models: The MCMCglmm R package. J. Stat. Softw. , 33,
1–22.
Hadfield, J.D., Wilson, A.J., Garant, D., Sheldon, B.C. & Kruuk, L.E.B.
(2010). The misuse of BLUP in ecology and evolution. Am. Nat. ,
175, 116–125.
Heidelberger, P. & Welch, P.D. (1983). Simulation Run Length Control in
the Presence of an Initial Transient. Oper. Res. , 31, 1109–1144.
Houslay, T.M. & Wilson, A.J. (2017). Avoiding the misuse of BLUP in
behavioural ecology. Behav. Ecol. , 28, 948–952.
Integrated-Informatics. (2014). Sustainable Development and Strategic
Science Branch Land Cover Classification.
Kasozi, H. & Montgomery, R.A. (2020). Variability in the estimation of
ungulate group sizes complicates ecological inference. Ecol.
Evol. , 1–9.
Laskowski, K.L. & Bell, A.M. (2013). Competition avoidance drives
individual differences in response to a changing food resource in
sticklebacks. Ecol. Lett. , 16, 746–753.
Leclerc, M., Vander Wal, E., Zedrosser, A., Swenson, J.E., Kindberg, J.
& Pelletier, F. (2016). Quantifying consistent individual differences
in habitat selection. Oecologia , 180, 697–705.
Lesmerises, F., Johnson, C.J. & St-Laurent, M.-H. (2018). Landscape
knowledge is an important driver of the fission dynamics of an alpine
ungulate. Anim. Behav. , 140, 39–47.
Mahoney, S.P., Lewis, K.P., Weir, J.N., Morrison, S.F., Luther, G.J.,
Schaefer, J.A., et al. (2016). Woodland caribou calf mortality in
Newfoundland: insights into the role of climate, predation and
population density over three decades of study. Popul. Ecol. , 58,
91–103.
Mahoney, S.P., Virgl, J.A., Fong, D.W., MacCharles, A.M. & McGrath, M.
(1998). Evaluation of a Mark-Resighting Technique for Woodland Caribou
in Newfoundland. J. Wildl. Manage. , 62, 1227–1235.
Mathot, K.J., Wright, J., Kempenaers, B. & Dingemanse, N.J. (2012).
Adaptive strategies for managing uncertainty may explain
personality-related differences in behavioural plasticity. Oikos ,
121, 1009–1020.
McLoughlin, P.D., Boyce, M.S., Coulson, T. & Clutton-Brock, T. (2006).
Lifetime reproductive success and density-dependent, multi-variable
resource selection. Proc. R. Soc. B , 273, 1449–1454.
McLoughlin, P.D., Morris, D.W., Fortin, D., Vander Wal, E. & Contasti,
A.L. (2010). Considering ecological dynamics in resource selection
functions. J. Anim. Ecol. , 79, 4–12.
Morris, D.W. (1987). Tests of density-dependent habitat selection in a
patchy environment. Ecol. Monogr. , 57, 269–281.
Morris, D.W. (1989). Density-dependent habitat selection: testing the
theory with fitness data. Evol. Ecol. , 3, 80–94.
Morris, D.W. (2003). Toward an ecological synthesis: A case for habitat
selection. Oecologia , 136, 1–13.
Newsome, S.D., Tinker, M.T., Gill, V.A., Hoyt, Z.N., Doroff, A., Nichol,
L., et al. (2015). The interaction of intraspecific competition
and habitat on individual diet specialization: a near range-wide
examination of sea otters. Oecologia , 178, 45–59.
Nicolaus, M., Tinbergen, J.M., Ubels, R., Both, C. & Dingemanse, N.J.
(2016). Density fluctuations represent a key process maintaining
personality variation in a wild passerine bird. Ecol. Lett. , 19,
478–486.
O’Brien, P.P., Webber, Q.M.R. & Vander Wal, E. (2018). Consistent
individual differences and population plasticity in network-derived
sociality: An experimental manipulation of density in a gregarious
ungulate. PLoS One , 13, e0193425.
Peignier, M., Webber, Q.M.R., Koen, E.L., Laforge, M.P., Robitaille,
A.L. & Vander Wal, E. (2019). Space use and social association in a
gregarious ungulate: Testing the conspecific attraction and resource
dispersion hypotheses. Ecol. Evol. , 9, 5133–5145.
Pelletier, F., Garant, D. & Hendry, A.P. (2009). Eco-evolutionary
dynamics. Philos. Trans. R. Soc. B , 364, 1483–1489.
Pyke, G.H., Pulliam, H.R. & Charnov, E.L. (1977). Optimal foraging: a
selective review of theory and tests. Q. Rev. Biol. , 52,
137–154.
R Core Team. (2020). R: A language and environment for statistical
computing. R Foundation for Statistical Computing, Vienna, Austria.
Robitaille, A.L., Webber, Q.M.R. & Vander Wal, E. (2019). Conducting
social network analysis with animal telemetry data: applications and
methods using spatsoc. Methods Ecol. Evol. , 10, 1203–1211.
Schaefer, J.A. & Mahoney, S.P. (2013). Spatial dynamics of the rise and
fall of caribou (Rangifer tarandus ) in Newfoundland. Can.
J. Zool. , 91, 767–774.
Sheppard, C.E., Inger, R., Robbie, A., Barker, S. & Emma, I.K. (2018).
Intragroup competition predicts individual foraging specialisation in a
group-living mammal. Ecol. Lett. , 21, 665–673.
Shizuka, D. & Johnson, A.E. (2020). How demographic processes shape
animal social networks. Behav. Ecol. , 31, 1–11.
Sikes, R.S. & Mammalogists, A.C. and U.C. of the A.S.O. (2016). 2016
Guidelines of the American Society of Mammalogists for the use of wild
mammals in research and education. J. Mammal. , 97, 663–688.
Smith, B.R. & Blumstein, D.T. (2008). Fitness consequences of
personality: A meta-analysis. Behav. Ecol. , 19, 448–455.
Stamps, J.A. (2016). Individual differences in behavioural plasticities.Biol. Rev. , 91, 534–567.
Strickland, K., Mann, J., Foroughirad, V., Levengood, A.L. & Frère,
C.H. (2021). Maternal effects and fitness consequences of individual
variation in bottlenose dolphins’ ecological niche. J. Anim.
Ecol. , 1948–1960.
Stuber, E.F., Carlson, B.S. & Jesmer, B.R. (2022). Spatial
personalities: a meta-analysis of consistent individual differences in
spatial behavior. Behav. Ecol. , 1–10.
Svanbäck, R. & Bolnick, D.I. (2007). Intraspecific competition drives
increased resource use diversity within a natural population.Proc. R. Soc. B , 274, 839–844.
Svanbäck, R. & Eklöv, P. (2006). Genetic variation and phenotypic
plasticity: causes of morphological and dietary variation in Eurasian
perch. Evol. Ecol. Res. , 37–49.
Tinker, M.T., Bentall, G. & Estes, J.A. (2008). Food limitation leads
to behavioral diversification and dietary specialization in sea otters.Proc. Natl. Acad. Sci. U. S. A. , 105, 560–565.
Vander Wal, E. & Webber, Q.M.R. (2020). Density dependence and
eco-evolutionary dynamics of animal social networks: a comment on
Shizuka and Johnson. Behav. Ecol. , 31, 16–18.
Vander Wal, E., Webber, Q.M.R. & Laforge, M.P. (2022). Repeatability is
the first step in a broader hypothesis test: a comment on Stuber et al.Behav. Ecol. , 1–2.
Webber, Q.M.R. & Vander Wal, E. (2018). An evolutionary framework
outlining the integration of individual social and spatial ecology.J. Anim. Ecol. , 87, 113–127.
Webber, Q.M.R. & Vander Wal, E. (2021). Context-dependent group size:
effects of population density, habitat, and season. Behav. Ecol. ,
1–12.
Wilson, A.J., Réale, D., Clements, M.N., Morrissey, M.M., Postma, E.,
Walling, C.A., et al. (2010). An ecologist’s guide to the animal
model. J. Anim. Ecol. , 79, 13–26.
Zaccarelli, N., Bolnick, D.I. & Mancinelli, G. (2013). RInSp: an R
package for the analysis of individual specialization in resource use.Methods Ecol. Evol. , 4, 1018–1023.