Author contributions
Yi-Hui Zhang and Hai-Lei Zheng designed the experiments. Zhi-Jun Shen, Kabir Ghoto, Gui-Feng Gao, Peng-Fei Li, Mei-Rong Luo, Zan Li, Zhi-Zhu Zhao and Yi-Hui Zhang performed the experiments. Zhi-Jun Shen, Kabir Ghoto and Yi-Hui Zhang analyzed the data. Zhi-Jun Shen and Yi-Hui Zhang wrote the paper. Yi-Hui Zhang and Hai-Lei Zheng revised this paper. All authors have read and approved the manuscript.