Acknowledgements
This project was supported by the National Key Research and Development Program of China (2016YFC0500602), the National Natural Science Foundation of China (31971744, 31770520), the State Basic Research Program (2012FY111900), the Program for Introducing Talents to Universities (B16011), and the German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig, the German Research Foundation (FZT 118), as well as by the European Research Council (ERC) under the European Union’s Horizon 2020 research and innovation program (grant agreement no 677232).