Acknowledgements
The following people provided help with logistics during fieldwork and/or labwork: . B. Alvarez, M. Ammer, Bahruddin, , H. Breeuwer, L. Dong, M. Erdmann, B. Hoeksema, P. Kuperus, S.C. Lim, S. Menken, A. Miners, A. Oherenan, K. Peijnenburg, W. Renema, Suharsono, R. Tapilatu, Y. Tuti, B. Voetdijk, N.J. de Voogd, the staff of Nabucco Island Dive Resort, Derawan Dive Resort, Misool Eco resort, Papua Diving. We are grateful for useful discussions with Dr. P. Meirmans. This work used the Vincent J. Coates Genomics Sequencing Laboratory at UC Berkeley, supported by NIH S10 Instrumentation grants S10RR029668 and S10RR027303. The Netherlands Organisation for Scientific Research provided funding to Leontine E. Becking through the grants RUBICON #825.12.007 and VENI #863.14.020. Fieldwork in Indonesia was made possible through additional financial support of National Geographic Society Waitt Grant, De Beukelaar-van der Hucht Stichting, World Wildlife Fund, Netherlands-INNO Fund, the Schure-Beijerinck-Popping Fund of the Royal Dutch Academy of Science (KNAW), Conservation International Ecosystem Based Management program (funded by the David and Lucile Packard Foundation), the Treub- Maatschappij Fund, the Leiden University Fund (LUF)/Slingelands, Singapore Airlines, the A.M. Buitendijk Fund and the J.J. ter Pelkwijk Fund (Naturalis Biodiveristy Center). The funders had no role in the study design, data collection and analysis, decision to publish, or preparation of the manuscript. We are grateful to the Indonesian Institute of Sciences (LIPI) and the Indonesian State Ministry of Research and Technology (RISTEK) for providing the research permits (0094/FRP/SM/V/2009;098/SIP/ FRP/SM/V/2011;098/SIP/FRP/SM/V/20113246/FRP/SM/VII/2012;#3B/TKPIPA/E5/Dit.KI/III/2016).