Author contributions
EB designed the study, sampled in the field, conducted molecular laboratory work, bioinformatics, analyzed molecular data, and drafted the manuscript. SB conducted chemistry laboratory work, analyzed chemistry data, and gave feedback on the manuscript. AN sampled in the field and gave feedback on the manuscript. SC provided input on the study and feedback on the manuscript. FJAN designed the study, helped with data analyses, and gave feedback on the manuscript. All authors gave final approval for publication.