creating pockets of clean fluid intermixed with fluid bearing odor of different concentrations (Campbell, 1977; Weissburg, 2000). These pockets of odor decrease in concentration as they are carried away from the source (Murlis and Jones, 1981; Murlis et al., 2000; Webster and Weissburg, 2001). In either flow condition, an animal must be able to adapt its behavior as it moves and use the information available to navigate to the source.